Non-Lepidopterous Invertebrates at Rough Bank
A scarce bug: Tettigometra impressopunctata
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Meloe rugosus Rugged Oil Beetle
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Bombylius discolor Dotted Bee-fly
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A scarce pot beetle: Cryptocephalus bipunctatus
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Whilst a lot of effort has been put into recording moths and butterflies at Rough Bank, relatively little
is known about all the other invertebrates on the reserve. This page lists
315 non-lepidopterous species which have been recorded here,
no doubt only a small fraction of the site's actual total which could be several thousand.
For Lepidoptera, the best-recorded large group of insects at the reserve, more than 25% of the
UK species (residents & frequent migrants) have been found here.
When managing for biodiversity, whilst it should be good to improve and increase the habitat for 'special' species
we know about, it is arguably much more important to avoid accidentally destroying the habitat for some of the
greater number of species of whose presence we are unaware!
This page lists many groups of terrestrial and freshwater invertebrates, most if not all of which include species which are present
on the reserve whether or not we have records of any of them. It does not include numerous groups which could be represented on
the reserve but which are less likely to be recorded, for example flukes, tapeworms and microscopic parasites, and
some aquatic groups.
Within each Class, Order or Family, species are listed in alphabetical order of scientific name.
Please note: groupings and names might not be up to date with changes in taxonomy and nomenclature.
Where families are listed under an order, only those families with species records are shown.
All records are from 2012 or later, and most have been made by members of the Gloucestershire Invertebrate Group.
Where numbers are shown in brackets, these are known/approximate/estimated numbers of British species, which in some cases
could include not just residents and frequent migrants but also rare migrants and accidentally imported species etc.
There are various sources for this data, providing varying totals for some species groups.
Phylum Arthropoda
Arthropods: Invertebrates with external skeletons, segmented bodies and jointed legs/appendages.
- Class Arachnida: Spiders & other Arachnids
- Order Araneae: Spiders (640)
- Family Theridiidae
- Family Linyphiidae
- Dismodiscus bifrons
- Entelecara acuminata
- Neriene peltata
- Porrhomma pygmaeum
- Tenuiphantes tenuis
- Family Araneidae
- Araneus quadratus
Four-spotted Orb-weaver
- Cercidia prominens
- Family Lycosidae
- Family Pisauridae
- Pisaura mirabilis
Nursery Web Spider
- Family Zoridae
- Family Thomisidae
Order Opiliones: Harvestmen (30)
- Homalenotus quadridentatus
- Mitostoma chrysomeles
- Nemastoma bimaculatum
- Oligolophus tridens
Order Pseudoscorpiones: Pseudoscorpions (27)
Order Trombidiformes: Mites
- Aceria macrochela
(galls on field maple)
Aceria myriadeum (galls on field maple)
Eriophyes similis (galls on blackthorn)
Phyllocoptes goniothorax (galls on hawthorn)
Order Parasitiformes: Ticks & Mites
- Ixodes ricinus
so beware when visiting!
Mites are also present, species undetermined
Class Branchiopoda: (aquatic crustaceans)
No species records
Class Malacostraca: (aquatic & terrestrial crustaceans)
- Order Bathynellacea: (groundwater crustaceans)
No species records
- Order Amphipoda: Amphipods
No species records
- Order Isopoda: Woodlice & Waterlice (60)
- Armadillidium vulgare
- Oniscus asellus
- Philoscia muscorum
- Platyarthrus hoffmannseggi
- Porcellio scaber
- Porcellio spinicornis
- Trichoniscus provisorius
Class Maxillopoda: (Copepods & other crustaceans)
No species records
Class Ostracoda: (aquatic crustaceans)
No species records
Class Insecta: Insects
- Order Zygentoma: Silverfish & Firebrat (2)
No species records
- Order Microcoryphia: Bristletails (7)
No species records
- Order Ephemeroptera: Mayflies (51)
No species records
- Order Odonata: Dragonflies & Damselflies (57)
- Family Coenagrionidae: Damselflies
- Coenagrion puella
Azure Damselfly
- Pyrrhosoma nymphula Large Red Damselfly
- Family Aeshnidae: Hawker Dragonflies
- Aeshna grandis
Brown Hawker
Family Libellulidae: Chasers, Skimmers & Darters
- Libellula depressa
Broad-bodied Chaser
Sympetrum striolatum Common Darter
Order Plecoptera: Stoneflies (34)
No species records
Order Orthoptera: Grasshoppers & Crickets (34)
- Family Tettigoniidae: Bush-crickets
- Meconema thalassinum
Oak Bush-cricket
- Metrioptera roeselii Roesel's Bush-cricket
- Pholidoptera griseoaptera Dark Bush-cricket
Family Tetrigidae: Ground-hoppers
- Tetrix undulata
Common Ground-hopper
Family Acrididae: Grasshoppers
- Chorthippus parallelus
Meadow Grasshopper
Gomphocerippus rufus Rufous Grasshopper
Stenobothrus lineatus Stripe-winged Grasshopper
Order Dermaptera: Earwigs (7)
- Forficula auricularia
Common Earwig
Order Blattodea: Cockroaches (3)
No species records
Order Psocodea: Booklice, Barklice & Parasitic Lice (650)
- Family Caeciliusidae: Barklice
- Family Ectopsocidae: Barklice
- Ectopsocus briggsi
- Ectopsocus petersi
Order Thysanoptera: Thrips (150)
No species records
Order Hemiptera: True Bugs (1800)
- Family Acanthosomatidae: Keeled Shieldbugs
- Acanthosoma haemorrhoidale
Hawthorn Shieldbug
Family Scutelleridae: Tortoise Shieldbugs
- Eurygaster testudinaria
Tortoise Shieldbug
Family Pentatomidae: Shieldbugs
- Aelia acuminata
Bishop's Mitre Shieldbug
Dolycoris baccarum Hairy Shieldbug
Eurydema oleracea Crucifer Shieldbug
Palomena prasina Green Shieldbug
Family Rhopalidae: Rhopalid Bugs
Family Berytidae: Stiltbugs
Family Nabidae: Damsel Bugs
- Himacerus apterus
Tree Damsel Bug
Family Anthocoridae: Flower Bugs
- Anthocoris nemorum
Common Flower Bug
Family Miridae: Plant Bugs
- Calocoris roseomaculatus
- Leptopterna dolabrata
Meadow Plant Bug
Megaloceroea recricornis
Phytocoris varipes
Stenodema calcarata
Stenodema laevigata
Family Gerridae: Pondskaters
- Gerris lacustris
Common Pondskater
Family Membracidae: Treehoppers
- Centrotus cornutus
Horned Treehopper
Family Cercopidae: Froghoppers
- Cercopis vulnerata
Red-and-Black Froghopper
Family Aphrophoridae: Froghoppers
- Aphrophora alni
Alder Spittlebug
Family Cicadellidae: Leafhoppers
- Alebra albostriella
- Cicadella viridis
- Dikraneura variata
- Edwardsiana flavescens
- Empoasca vitis
- Mocydia crocea
- Planaphrodes bifasciata
- Zyginidia scutellaris
Family Delphacidae: Planthoppers
Family Tettigometridae: Planthoppers
- Tettigometra impressopunctata
Family Aphididae: Aphids
- Dysaphis ranunculi
(gall on hawthorn)
Family Psyllidae: Jumping Plant Lice
Family Triozidae: Jumping Plant Lice
- Trioza remota
- Trioza urticae
Order Neuroptera: Lacewings (68)
Order Megaloptera: Alderflies (3)
No species records
Order Raphidioptera: Snakeflies (4)
- Atlantoraphidia maculicollis
Order Coleoptera: Beetles (4000)
- Family Carabidae: Tiger & Ground Beetles
- Abax parallelepipedus
- Demetrias atricapillus
- Harpalus latus
- Harpalus rufipes
- Nebria brevicollis
- Paradromius linearis
- Pterostichus madidus
Black Clock Beetle
- Pterostichus melanarius
- Stomis pumicatus
Family Silphidae: Burying Beetles
- Necrodes littorialis
- Nicrophorus humator
Black Sexton Beetle
Nicrophorus vespillo
Silpha atrata Black Snail Beetle
Family Staphylinidae: Rove Beetles
- Anotylus sculpturatus
- Drusilla canaliculata
- Habrocerus capillaricornis
- Staphylinus dimidiaticornis
- Stenus aceris
Family Geotrupidae: Dor Beetles
- Geotrupes spiniger
Common Dumble Dor
Family Lucanidae: Stag Beetles
- Sinodendron cylindricum
Rhinoceros Beetle
Family Scarabaeidae: Scarab Beetles
- Aphodius pedellus
- Aphodius sphacelatus
- Melolontha melolontha
Cockchafer
Phyllopertha horticola Garden Chafer
Family Dascillidae: Soft-bodied Plant Beetles
- Dascillus cervinus
Orchid Beetle
Family Elateridae: Click Beetles
- Athous haemorrhoidalis
- Limonius minutus
(split to L. poneli in some references)
Prosternon tessellatum Chequered Click Beetle
Family Lampyridae: Fireflies
- Lampyris noctiluca
Glow-worm
Family Cantharidae: Soldier Beetles
- Cantharis pellucida
- Cantharis rufa
- Cantharis rustica
- Rhagonycha fulva
Common Red Soldier Beetle
Rhagonycha lignosa
Family Malachiidae: Soft-winged Flower Beetles
- Malachius bipustulatus
Common Malachite
Family Coccinellidae: Ladybirds
- Chilocorus renipustulatus
Kidney-spot Ladybird
Coccinella septempunctata 7-spot Ladybird
Halyzia sedecimguttata Orange Ladybird
Harmonia axyridis Harlequin Ladybird
Propylea quattuordecimpunctata 14-spot Ladybird
Subcoccinella vigintiquattuorpunctata 24-spot Ladybird
Family Mycetophagidae: Hairy Fungus Beetles
- Mycetophagus quadripustulatus
Family Oedemeridae: False Blister Beetles
- Ischnomera cyanea
- Oedemera femoralis
- Oedemera lurida
- Oedemera nobilis
Swollen-thighed Beetle
Family Meloidae: Oil Beetles
- Meloe rugosus
Rugged Oil Beetle
Family Pyrochroidae: Cardinal Beetles
- Pyrochroa serraticornis
Red-headed Cardinal Beetle
Family Salpingidae: Narrow-waisted Bark Beetles
Family Cerambycidae: Longhorn Beetles
- Rhagium mordax
Black-spotted Longhorn Beetle
Stenocorus meridianus Variable Longhorn Beetle
Stenurella melanura Black-striped Longhorn Beetle
Family Chrysomelidae: Leaf Beetles
- Altica helianthemi
- Aphthona euphorbiae
- Cassida rubiginosa
Thistle Tortoise Beetle
Cryptocephalus aureolus
Cryptocephalus bipunctatus
Cryptocephalus hypochaeridis
Gastrophysa viridula Green Dock Beetle
Lochmaea crataegi Hawthorn Leaf Beetle
Longitarsus suturellus
Mantura matthewsii
Neocrepidodera ferruginea
Oulema melanopus Cereal Leaf Beetle
Oulema obscura
Pyrrhalta viburni Viburnum Leaf Beetle
Timarcha goettingensis Small Bloody-nosed Beetle
Family Anthribidae: Weevils
- Platyrhinus resinosus
Cramp-ball Fungus Weevil
Platystomos albinus Bird-dropping Weevil
Family Attelabidae: Leaf-roller Weevils
- Apoderus coryli
Hazel Leaf-roller Weevil
Family Apionidae: Weevils
Family Curculionidae: Weevils
- Cionus hortulanus
- Rhamphus oxyacanthae
(leaf-mine on hawthorn)
Order Strepsiptera: Twisted-wing Parasites (10)
No species records
Order Mecoptera: Scorpionflies (4)
- Panorpa cognata
Scarce Scorpionfly
Panorpa germanica Saxon Scorpionfly
Order Trichoptera: Caddisflies (200)
No species records
Order Lepidoptera: Moths & Butterflies (2500)
More than 760 species recorded at the reserve
For species lists, see the
Moths (full list) and
Butterflies pages
Order Diptera: Flies (7000)
- Family Tipulidae: Craneflies
- Tipula maxima
Giant Cranefly
Family Cecidomyiidae: Gall Midges
- Contarinia coryli
(gall & larvae on hazel)
Hartigiola annulipes (gall on beech)
Family Psychodidae: Moth Flies
- Boreoclytocerus ocellaris
- Pericoma pilularia
- Philosepedon humeralis
- Psychoda albipennis
- Psychoda grisescens
- Psychoda phalaenoides
Family Ptychopteridae: Fold-winged Craneflies
Family Rhagionidae: Snipeflies
- Chrysopilus asiliformis
Little Snipefly
Family Tabanidae: Horseflies
- Haematopota pluvialis
Notch-horned Cleg
Family Bombyliidae: Bee-flies
- Bombylius discolor
Dotted Bee-fly
Bombylius major Dark-edged Bee-fly
Villa cingulata Downland Villa
Family Asilidae: Robberflies
- Leptogaster cylindrica
Slender-striped Robberfly
Machimus atricapillus Kite-tailed Robberfly
Family Syrphidae: Hoverflies
- Cheilosia albitarsis
- Cheilosia vernalis
- Chrysotoxum bicinctum
- Eristalis arbustorum
- Eristalis nemorum
- Eristalis pertinax
- Eristalis tenax
- Eupeodes corollae
- Ferdinandea cuprea
- Helophilus pendulus
- Helophilus trivittatus
- Melanostoma mellinum
- Melanostoma scalare
- Neoascia podagrica
- Platycheirus albimanus
- Platycheirus clypeatus
- Rhingia campestris
- Scaeva pyrastri
- Sericomyia superbiens
- Syrphus vitripennis
- Volucella inflata
Family Conopidae: Thick-headed Flies
Family Ulidiidae: Picture-winged Flies
Family Tephritidae: Picture-winged Flies
- Acidia cognata
(mine on coltsfoot)
Tephritis formosa
Tephritis vespertina
Urophora cardui (gall on creeping thistle)
Family Sciomyzidae: Snail-killing Flies
Family Sepsidae: Ensign Flies
Family Agromyzidae: Leaf-miner Flies
- Amauromyza flavifrons
(mine on red campion)
Chromatomyia scolopendri (mine on hartstongue)
Liriomyza congesta (mine on horseshoe vetch)
Phytobia cerasiferae (mine in blackthorn stem)
Phytomyza glechomae (mine on ground ivy)
Phytomyza ilicis (mine on holly)
Phytomyza lappae (mine on burdock)
Phytomyza minuscula (mine on columbine)
Phytomyza ranunculi (mine on lesser celandine)
Phytomyza tussilaginis (mine on coltsfoot)
Family Muscidae: includes House Flies
Family Calliphoridae: Blowflies
- Pollenia rudis
Cluster Fly
Order Siphonaptera: Fleas (60)
No species records
Order Hymenoptera: Ants, Bees, Wasps & Sawflies (7000)
- Family Cimbicidae: Sawflies
- Family Tenthredinidae: Sawflies
- Athalia rosae
Turnip sawfly
- Caliroa cerasi
- Fenella nigrita (leaf-mines on agrimony)
- Fenusa pumila (leaf-mine on birch)
Family Cynipidae: Gall-wasps
- Andricus lignicola
(gall on oak)
Diastrophus rubi (gall on bramble)
Diplolepis rosae (gall on rose)
Neuroterus numismalis (gall on oak)
Neuroterus quercusbaccarum (gall on oak)
Family Ichneumonidae: Ichneumon Wasps
- Amblyteles armatorius
- Ichneumon sarcitorius
- Netelia inedita
- Ophion obscuratus
Family Formicidae: Ants
- Formica fusca
- Lasius alienus
- Lasius flavus
Yellow Meadow Ant
Lasius niger Black Garden Ant
Myrmica sabuleti
Myrmica scabrinodis
Family Vespidae: Social, Potter & Mason Wasps
- Dolichovespula sylvestris
Tree Wasp
Vespa crabro Hornet
Vespula germanica German Wasp
Vespula vulgaris Common Wasp
Family Crabronidae: Digger Wasps
Family Andrenidae: Mining Bees
- Andrena bucephala
Big-headed Mining Bee
Andrena cineraria Ashy Mining Bee
Andrena fulvago Hawk's-beard Mining Bee
Andrena haemorrhoa Early Mining Bee
Andrena scotica Chocolate Mining Bee
Andrena trimmerana Trimmer's Mining Bee
Andrena wilkella Wilke's Mining Bee
Family Halictidae: Furrow Bees
- Halictus tumulorum
Bronze Furrow Bee
Lasioglossum calceatum Common Furrow Bee
Lasioglossum fulvicorne Chalk Furrow Bee
Lasioglossum morio Green Furrow Bee
Sphecodes hyalinatus Furry-bellied Blood Bee
Sphecodes monilicornis Box-headed Blood Bee
Family Megachilidae: includes Mason Bees
- Osmia bicolor
Red-tailed Mason Bee
Osmia spinulosa Spined Mason Bee
Family Apidae: includes Bumblebees and Nomad Bees
- Bombus hypnorum
Tree Bumblebee
Bombus lapidarius Red-tailed Bumblebee
Bombus lucorum White-tailed Bumblebee
Bombus pascuorum Common Carder Bee
Nomada flavoguttata Little Nomad Bee
Nomada hirtipes Long-horned Nomad Bee
Nomada lathburiana Lathbury's Nomad Bee
Class Entognatha: (6-legged wingless arthropods)
- Order Diplura: Two-pronged Bristletails (11)
No species records
- Order Protura: Coneheads (15)
No species records
- Order Collembola: Springtails (250)
- Family Entomobryidae
- Lepidocyrtus cyaneus
- Lepidocyrtus lanuginosus
- Orchesella cincta
- Orchesella villosa
- Family Isotomidae
- Anurophorus laricis
- Pseudisotoma sensibilis
- Family Tomoceridae
- Family Katiannidae
Class Chilopoda: Centipedes (59)
- Lithobius calcaratus
- Lithobius microps
Class Diplopoda: Millipedes (63)
- Cylindroiulus caeruleocinctus
- Glomeris marginatus
- Ophyiulus pilosus
- Tachypodoiulus niger
Class Symphyla: Pseudocentipedes (14)
No species records
Class Pauropoda: Pauropods (23)
No species records
Phylum Annelida: Ringed Worms
- Class Oligochaeta: (includes Earthworms (26))
No species records
Phylum Mollusca: Molluscs
- Class Gastropoda: Snails & Slugs (190)
- Family Pomatiidae: Snails
- Family Cochlicopidae: Snails
- Cochlicopa lubrica
- Cochlicopa lubricella
- Family Pyramidulidae: Snails
- Family Chondrinidae: Snails
- Family Lauriidae: Snails
- Family Valloniidae: Snails
- Family Enidae: Snails
- Ena montana
- Merdigera obscura
- Family Discidae: Snails
- Family Arionidae: Slugs
- Arion ater
- Arion circumscriptus
- Arion distinctus
- Arion rufus
- Arion silvaticus
- Arion subfuscus
- Family Oxychilidae: Glass Snails
- Aegopinella nitidula
- Aegopinella pura
- Oxychilus cellarius
- Family Vitrinidae: Snails
- Family Boettgerillidae: Slugs
- Family Limacidae: Keelback Slugs
- Lehmannia marginata
- Limax maximus
- Family Agriolimacidae: Slugs
- Deroceras invadens
- Deroceras reticulatum
- Family Ferussaciidae: Snails
- Family Clausiliidae: Door Snails
- Balea heydeni
- Clausilia bidentata
- Cochlodina laminata
- Family Hygromiidae: Snails
- Candidula intersecta
- Helicella itala
- Trochulus hispidus
- Trochulus striolatus
- Zenobiella subrufescens
- Family Helicidae: Snails
- Arianta arbustorum
- Cepaea hortensis
- Cepaea nemoralis
- Cornu aspersum
- Helicigona lapicida
- Helix pomatia
Roman Snail